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Darwin, Charles, 1809-1882

"On the Origin of Species By Means of Natural Selection, or, the Preservation of Favoured Races in the Struggle for Life"


With pigeons, however, we have another case, namely, the occasional
appearance in all the breeds, of slaty-blue birds with two black bars
on the wings, a white rump, a bar at the end of the tail, with the
outer feathers externally edged near their bases with white. As all
these marks are characteristic of the parent rock-pigeon, I presume
that no one will doubt that this is a case of reversion, and not of a
new yet analogous variation appearing in the several breeds. We may I
think confidently come to this conclusion, because, as we have seen,
these coloured marks are eminently liable to appear in the crossed
offspring of two distinct and differently coloured breeds; and in this
case there is nothing in the external conditions of life to cause the
reappearance of the slaty-blue, with the several marks, beyond the
influence of the mere act of crossing on the laws of inheritance.
No doubt it is a very surprising fact that characters should reappear
after having been lost for many, perhaps for hundreds of generations.
But when a breed has been crossed only once by some other breed, the
offspring occasionally show a tendency to revert in character to the
foreign breed for many generations--some say, for a dozen or even a
score of generations. After twelve generations, the proportion of
blood, to use a common expression, of any one ancestor, is only 1 in
2048; and yet, as we see, it is generally believed that a tendency to
reversion is retained by this very small proportion of foreign blood.


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